Presumably there is some premailer or inline-styler used to build template/default.tmpl's email.default.html value from template/email.html. However, from a cursory glance, it appears out of sync on #1475. Specifically that PR didn't update the default template HTML.
Are they out of sync by intention? What tool is used to generate the inline-style versioned? I came across this when working with my own templates.